CARMIL1 (Capping Protein Regulator and Myosin 1 Linker 1): CARMIL1 is a gene that encodes a protein involved in regulating actin dynamics, a key part of the cellular cytoskeleton. It plays an important role in cell movement, structure, and signaling. Changes or disruptions in CARMIL1 can impact cell motility and have been associated with immune system dysfunction and certain cancers.